Recalcitrant seeds are seeds that do not survive drying and freezing during ex situ conservation. By and large, these seeds cannot resist the effects of drying or temperatures less than 10 °C ; thus, they cannot be stored for long periods like orthodox seeds because they can lose their viability. Plants that produce recalcitrant seeds include avocado, mango, mangosteen, lychee, cocoa, rubber tree, some horticultural trees, aquatic plants such as Nymphaea caerulea, and several plants used in traditional medicine, such as species of Virola and Pentaclethra. Generally speaking, most tropical pioneer species have orthodox seeds but many climax species have recalcitrant or intermediate seeds.